The China-born Dr Cai, who became a Singapore citizen, obtained a PhD in molecular biology from Stanford University in 1990. The Straits Times learnt that he did a two-year postdoctoral fellowship after leaving Stanford at the University of Washington, under famed genetist Professor Lee Hartwell, who won a 2001 Nobel Prize in physiology.

He joined IMCB two years later and worked as a principal investigator in the field of cell genetics up till his departure